Alex the lion (Ben Stiller), Marty the zebra (Chris Rock), giraffe Melman (David Schwimmer) and hippopotamus Gloria (Jada Pinkett Smith) and friends decide to escape from Madagascar. But by coincidence all the company find themselves in Africa, where Alex meets his family, with whom he was separated in childhood. Unexpectedly Alex understands that after life in the zoo he had much less to do with family than he would like to. Meanwhile Gloria starts on to finding the man of her life, but discovers that sometimes love seems further than it is...